1904 Nobel Laureate in Medicine

    in recognition of his work on the physiology of digestion, through which knowledge on vital aspects of the subject has been transformed and enlarged.

  • September 14, 1849 - February 27, 1936
  • Place of birth: Ryazan (Russia)
  • Education: Studied medicine at St. Petersburg (Russia)
  • Affiliation: Institute of Experimental Medicine, St. Petersburg (Professor from 1890; Director from 1902); Military Medical Academy, St. Petersburg
  • Worked on the physiology of circulation and digestion
  • Definitive work: Lectures on Conditioned Reflexes (1926)
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